Evolutionarily, amphibians are somewhat simpler than mammals, they're smaller than a lot of mammals and they don't live as long, so I suspect some of this is simply that "things that aren't there". They wouldn't have as many problems with advanced glycation end products because the temperature is so low. There's at least one other ice survival strategy: antifreeze proteins. The fir tree and a variety of arctic fish have these: https://pmc.ncbi.nlm.nih.gov/articles/PMC6691018/.
In all cases, I still don't understand how the membrane potentials are maintained or re-constructed in the thawing phase: any pointers?
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